Asteraceae

Carduus hamulosus subsp. hamulosus

Carduus hamulosus subsp. hamulosus

Carduus L. Carduus hamulosus EHRH. subsp. hamulosus EHRH. Ömür: İki yıllık Yapı: ot Hayat formu: İlk çiçeklenme zamanı: 6 Son çiçeklenme zamanı: 8 Habitat: yüksek kesimler, çayırlık, otlak, nadas tarla, killi toprak, kumlu toprak Minimum yükseklik: 1210 Maksimum yükseklik: 2150 Endemik: - Element: Avrupa-Sibirya Türkiye dağılımı: D. Anadolu Genel dağılımı: D. Avusturya, Macaristan, G. Rusya, Kırım, Kafkasya, KB. İran Bulunduğu kareler:A8 A9 B9 B10

C. hamulosus Ehrh., Beitr. Naturk. 7:166 (1792). Biennial, 20-60(-100) cm, sparsely arachnoid. Stem narrowly undulate-winged with triangular, shortly spinulose-margined lobes, sparsely branched. Cauline leaves pinnatifid to ½-¾, primary segments 2-3-fid into ovate-triangular, shortly spinulose-margined lobes, often grey-tomentellous below. Peduncles long, 5-20 cm, spinulose-lobulate below, ± naked and tomentose above. Involucre solitary, depressed-globose, subglabrous, 20-30 x 25-40 mm; phyllaries gradually imbricate or outer ½-1 x as long as median, lax, outer lanceolate, median lanceolate-linear, 1-nerved above broadened base, ± recurved towards acute spinulose apex. Corollas zygomorphic, purple (rarely white), 18-25 mm. Achenes c. 4 mm, brown?; pappus 14-17 mm. Fl. 6-8. 1. Phyllaries attenuate above broadened basal portion, gradually enlarging from base to apex of involucre subsp. hamulosus 1. Phyllaries more gradually attenuate from near base, outer ½-1 x as long as median subsp. hystrix subsp. hamulosus. Syn: C. nigrescens Vill. subsp. hamulosus (Ehrh.) Arenes in Mem. Mus. Hist. Nat., Paris, n.s. 24:203 (1949); C. tauricus Klokov in Fl. URSR 11:569, t. 103 (1962); ? C. tyriacus Klokov, op. cit. 568, t. 102; C. pseudocollinus (Schmalh.) Klokov, op. cit. 510. Ic: Reichb., Ic. Fl. Germ. 15: t. 875 (1853); Jáv. & Csap., Ic. Fl. Hung. 538 (1953). Map 46, p. 433. Highlands, in meadows and pastures, fallow fields, on clay and sandy soils, 1210-2150 m. Type: Hungaria?, J.J. Winter 1. E. Anatolia. A8 Erzurum: 59 km from Erzurum to Ispir, 1900 m, Hub.-Mor. 15955! A9 Kars: Kars to Ardahan, 1790 m, 24 vi 1964, M.Zohary & Plitm! B9 Ağri: 32 km W. of Doğubayazit, 1210 m, Furse 3554! 3553 (fls. white)! B9 Van: Hoşap to W. foot of Kepir Da., 1850 m, D. 47238! B1O Van: 5 km N.E. of Başkale, D. 45879! E. Austria to Hungary, S. Russia,/Crimea, Caucasia; N.W. Iran? Euro-Sib. element. subsp. hystrix (C.A.Meyer) Kazmi in Mitt. Bot. Staatssamml. München 5:402 (1964). Syn: C. hystrix C.A.Meyer, Verz. PL Cauc. 71 (1831). Ic: Fl. URSS 28: t. 2 f. 3 (1963). Banks and fallow fields, 1700-1900 m. Type: in pratis siccis montium Talüsch (Talysh), 105-150 m, C.A.Meyer (LE, photo!). E. Anatolia. B9 Van: 5-8 km from Van to Değirmenköy, 1900 m, D. 44431! Ağri: 2 km S.W. of Hamur (Murat De.), 1700 m, D. 44176! Transcaucasia, N.W, Iran. Ir.-Tur. element. Although Turkish material matches the type of C. hystrix in the form of its involucre, the phyllaries are not arachnoid nor are the leaves densely tomentellous below. See remarks under C. uncinatus Bieb. on p. 438.